Skip to main content Skip to main content
Programs Overview
Undergraduate Programs
Graduate Programs
Course Catalog
Outreach Initiatives
Research Clusters
Research Areas
Funding Opportunities
People Overview
Find a Mentor
Alumni Directory
Job Opportunities
Admissions Overview
Undergraduate Admissions
Graduate Admissions
Contact Us
Maps & Directions
Campus Tours

Irv Kornfield

If you need to locate a faculty member with knowledge about particular functions of SMS (e.g., graduate admissions), you can find them on the list of current faculty committees.

Irv Kornfield

Contact Information

Irv Kornfield



215 Murray Hall
University of Maine 
Orono, ME  04469-5741


Ph.D. Stony Brook

Research interests

A principal thrust of current research employs molecular techniques to examine population processes and the systematics of marine organisms. We examine both small scale features of population structuring (genic isolation and gene flow) as well as large scale features (biogeographic reconstruction.) Animals studied include commercially important species (such as lobster and haddock) as well as widely distributed taxa (such as crabs, hemicordates, and cumaceans). This last group of organisms, a marine order of malacostracan crustaceans, is particularly interesting.  Systematic and population problems are examined via analysis of variation in DNA sequences and at microsatellite loci.   A second major area of research centers about speciation and radiation of fish species flocks with emphasis on cichlid fishes. Because of the complexity and rate of their radiation, African cichlids are available as a paradigm for evolutionary biology. Recent studies include characterizations of population processes and systematics.  


  • Netherland Journal Zoololgy
    McElroy, D.M., I. Kornfield and J. Everett. 1991. Coloration in African cichlids: diversity and constraints in Lake Malawi endemics. Neth. J. Zool. 41:250-268.
  • Proc. Biol. Soc. Washington
    King, G.M., C. Giray and I. Kornfield. 1994. A new hemicordate, Saccoglossus bromophenolicum from North America. Proc. Biol. Soc. Wash. 107:383-390.
  • Journal of Molecular Evolution
    Parker, A. and I. Kornfield. 1997. Evolution of the mitochondrial DNA control region in the mbuna (Cichlidae) species flock of Lake Malawi, East Africa. J. Mol. Evol. 45:70-83.
  • 2000 Molecular Ecology
    Brown, B.L., D.E. Franklin, P.M. Gaffney, M.Hong, D.DenDanto and I. Kornfield 2000. Characterization of microsatellite loci in the eastern oyster, Crassostrea virginica. Mol. Ecol. 9: 2217-2219.
  • Ann. Rev. Ecology
    Kornfield, I. and P.F. Smith. 2000. African cichlid fishes: model systems for evolutionary biology. Ann. Rev. Ecol. Syst. 31: 163-196.
  • Canadian Journal of Fisheries Aquaculture
    Lage, C., M.R. Purcell, and I. Kornfield. Genetic variation of haddock (Melanogrammus aeglefinnus) stocks in the Northwestern Atlatnic Ocean. Can. J. Fish. Aquat. Sci.(in press)
  • Molecular Ecology
    Lage, C.R. and I. Kornfield 1999. Isolation and characterization of microsattellite loci in Atlantic haddock (Melanogrammus aeglefinus). Mol. Ecol. 8: 1355-1357.
  • Journal of Crustacean Biology
    Y.K. Tam and I. Kornfield. 1998. Phylogenetic relationships among clawed lobster genera (Decapoda: Nephropidae) based on mitochondrial 16S rRNA gene sequence. J. Crust. Biol. 18:138-146.
  • Northeast Regional Agricultural Engineering Service
    Hunt von Herbing, I., I. Kornfield, M. Tupper and J. Wilson (eds.) 1998. The implications of Localized Fishery Stocks. Northeast Regional Agricultural Engineering Service, New York.
  • Academic Press
    Kornfield, I. and A. Parker. 1997. Molecular systematics of a rapidly evolving species flock: the mbuna of Lake Malawi and the search for phylogenetic signal. pp. Molecular systematics of fishes. Kocher, T.D. and C. Stepien (eds.). Academic Press, New York.
  • 1996 Marine Biology
    Tam, Y.K., I. Kornfield and F.P. Ojeda. 1996. Divergence and zoogeography of mole crabs, Emerita (Decapoda, Hippidae), in the Americas. Marine Biol. 125:489-498.
  • Genome
    Parker, A. and I. Kornfield. 1996. An improved amplification and sequencing strategy for phylogenetic studies using the mitochondrial large subunit rRNA gene. Genome. 39:793-797.
  • Molecular Marine Biology Betechnology
    Purcell, M.K., I. Kornfield, M.J. Fogarty, and A. Parker. 1996. Interdecadal heterogeneity of mitochondrial DNA in Atlantic haddock, Melanogrammus aeglefinus, from Georges Bank. Mol. Marine Biol. Biotech. 5:185-192.

    Tam, Y.K. and I. Kornfield. 1996. Characterization of microsatellite markers in Homarus (Crustacea, Decapoda). Mol. Marine Biol. Biotech. 5:230-238.
  • Environmenal Biological Fishes
    Parker, A. and I. Kornfield. 1996. Polygynandry in Pseudotropheus zebra, a cichlid fish from Lake Malawi. Envir. Biol. Fishes 47:345-352.
  • Fishery Bulletin
    Kornfield, I., A. Williams, and R.S. Steneck. 1995. Assignment of Homarus capensis (Herbst, 1792), the Cape lobster of South Africa, to Homarius new genus (Decapoda: Nephropidae). Fish. Bull. 93:97-102.
  • 1995 Copeia
    Parker, A. and I. Kornfield. 1995. A molecular perspective on the evolution and zoogeography of cyprinodontid killifishes (Teleostei, Atherinomorpha). Copeia 1995:8-21.
  • North American Journal of Fishery Biology
    Kircheis, F.W., I. Kornfield and S. Seyoum. 1995. The genetic identity of an introduced char. North Amer. J. Fish. Biol. 15:54-59.
  • 1995 Marine Biology
    King, G. M., C. Giray and I. Kornfield. 1995. Biogeographical, biochemical and genetic differentiation among North American saccoglossids (Hemichordata; Enteropneusta; Harrimaniidae). Marine Biol. 123:369-378.
  • Molecular Biology and Evolution
    Moran, P. and I. Kornfield. 1995. Were population bottlenecks associated with radiation of the mbuna species flock (Teleostei: Cichlidae) of Lake Malawi? Mol. Biol. Evol. 12:1085-1093.
  • Molecular Phylogenetic Evolution
    Franck, J.P.C., I. Kornfield and J.M. Wright. 1994. The utility of SATA satellite DNA sequences for inferring phylogenetic relationships among the three major genera of tilapiine cichlid fishes. Mol. Phylogenet. Evol. 3:10-16.
  • 1993 Copeia
    McElroy, D. and I. Kornfield. 1993. Hybridization and morphological differentiation between two Malawi cichlids. Copeia. 1993:933-939.
  • Aquaculture
    Seyoum, S. and I. Kornfield. 1992. Identification of the subspecies of Oreochromis niloticus (Pisces: Cichlidae) using restriction endonuclease analysis of mitochondrial DNA. Aquaculture 102:29-42.
  • Journal of Heredity
    McElroy, D. P. Moran, E. Birmingham and I. Kornfield. 1992. REAP: an intregrated program for the analysis of restriction fragment data. J. Heredity 83:157-158.
  • Canadian Journal of Zoology
    Seyoum, S. and I. Kornfield. 1992. Taxonomic notes on the Oreochromis niloticus subspecies complex (Pisces: Cichidae), with a description of a new subspecies. Can. J. Zool. 70:2161-2165.
  • Ecology and Evolution
    Kornfield, I. 1991. Genetics. pp. 103-128. in Cichlid Fishes: Behavior, Ecology and Evolution. Keenleyside, M. (ed.). Chapman and Hall.
  • 1990 Copeia
    Herke, S.W., I. Kornfield, P. Moran and J.R. Moring. 1990. Molecular confirmation of hybridization between northern pike (Esox lucius) and chain pickerel (E. niger). Copeia 1900:842-846.
  • 1990 Canadian Journal of Fishery Aquatic Science
    Stephenson, R.L., and I. Kornfield. 1990. Reappearance of spawning herring on Georges Bank: population resurgence not recolonization. Can. J. Fish. Aqu. Sci. 47:1060-1064.
  • Proc. R. Soc. Lond.
    Owen, R.B., R. Crossley, T.C. Johnson, D. Tweddle, I. Kornfield, S. Davison, D.H. Eccles and D.E. Engstrom. 1990. Major low levels of Lake Malawi and implication for speciation rates in cichlid fishes. Proc. R. Soc. Lond. 240B:519-553.
  • Environmental Biological Fishes
    McElroy, D.M. and I. Kornfield. 1990. Sexual selection, reproductive behavior, and speciation in the mbuna species flock of Lake Malawi (Pisces: Cichlidae). Envir. Biol. Fishes. 28:285-294.
  • American Science
    Kornfield, I. 1990. Community and Evolutionary Ecology of North American Stream Fishes. Amer. Sci. 78:64.
  • Journal of Food Comp.
    Bergeron, R.J. Bushway, F.L. Roberts, I. Kornfield, J. Okedi and A. Bushway. 1988. The nutrient composition of an insect flour from Lake Victoria, Uganda. J. Food Comp. Anal. 1: 371-377.
  • Fishery Bulletin
    Kornfield, I. and S.M. Bogdanowicz. 1987. Mitochondrial DNA heterogeneity in Atlantic herring. Fishery Bull. 85:561-568.
  • University of Maine at Orono Press
    Echelle, A.A. and I. Kornfield (eds.). 1984. Evolution of Fish Species Flocks. University of Maine at Orono Press.
  • 1984 Evolution
    McKaye, K.R., T. Kocher, P. Reinthal, R. Harrison and I. Kornfield. 1984. Genetic evidence for allopatric and sympatric differentiation among morphs of a Lake Malawi cichlid fish. Evolution 38:215-219.
  • Plenum Publication
    Kornfield, I. 1984. Descriptive genetics of cichlid fishes. pp. 591-616. in Evolutionary Genetics of Fishes. Turner, B.J. (ed.). Plenum Publ.
  • 1984ta University of Maine at Orono Press
    Kornfield, I. and K. Carpenter. 1984. The cyprinids of Lake Lanao, Philippines: taxonomic validity, evolutionary rates and speciation scenarios. pp. 69-84 in Evolution of Fish Species Flocks. Echelle, A.A. and I. Kornfield (eds.). University of Maine at Orono Press.
  • Proc. Biol. Soc. Wash.
    Kornfield, I. and J.N. Taylor. 1983. A new species of polymorphic fish, Cichlasoma minckleyi from Cuatro Cienegas, Mexico (Teleostei: Cichlidae). Proc. Biol. Soc. Wash. 96:253-269.
  • 1983 Quart. Rev. Biology
    Kornfield, I. 1983. The Haplochromine Fishes of the East African Lakes. Quart. Rev. Biol. 58:86.
  • 1982 Copeia
    Kornfield, I. 1982. Report from Mindanao. Copeia 1982:493-495.
  • Journal of Linn. Society
    McKaye, K.R., T. Kocher, P. Reinthal, and I. Kornfield. 1982. Sympatric sibling species of Petrotilapia Trewavas analyzed by enzyme electrophoresis (Pisces: Cichlidae). J. Linn. Soc. 76:91-96.
  • 1982 Evolution
    Kornfield, I.L., D.C. Smith, P.S. Gagnon and J.N. Taylor. 1982. The cichlid fish of Cuatro Cienegas, Mexico: direct evidence of conspecificity among distinct trophic morphs. Evolution 36:658-664.
  • Canadian Journal of Fisheries Aquatic science
    Kornfield, I.L., B.D. Sidell and P.S. Gagnon. 1982. Stock definition in Atlantic herring: genetic evidence for discrete fall and spring spawning populations. Can. J. Fish. Aqu. Sci. 39:1610-1621.
  • Canadian Journal of Fisheries Aquatics Science
    Kornfield, I.L., K.F. Beland, J.R. Moring and F.W. Kircheis. 1981. Genetic similarity of endemic arctic char (Salvelinus alpinus) and implications for their management. Can. J. Fish. Aqu. Sci. 38:32-39.
  • Copeia
    Kornfield, I.L. 1981. Distribution of constitutive heterochromatin and the evolution of sex chromosomes in Fundulus. Copeia 1981:916-918.
  • Progress Fish-Culture
    Smith, D.C. and I. Kornfield. 1981. A remotely operated trap for capture of territorial fishes. Prog. Fish. Cult. 43:208-209.
  • Canadian Journal of Genetics and Cytol.
    Kornfield, I., P.S. Gagnon and B.D. Sidell. 1981. Inheritance of allozymes in Atlantic herring (Clupea harengus harengus). Can. J. Genet. Cytol. 23:715-720.
  • 1979 Evolution
    Kornfield, I.L., U. Ritte, C. Richler and J. Wahrman. 1979. Biochemical and cytological aspects of species differentiation in Old World cichlid fishes. Evolution 33:1-14.
  • Experientia
    Kornfield, I.L. 1978. Evidence for rapid speciation in African cichlid fishes. Experientia 34:335-336.
  • Evol. Biological Fishes
    Kornfield, I. 1977. Investigation of the Ichthyofauna of Nicaraguan Lakes. Evol. Biol. Fishes 2:93-94.
  • Nature
    Kornfield, I.L. and E. Nevo. 1976. Likely pre-Suez occurrenceof a Red Sea Fish Aphanius dispar in the Mediterranean. Nature 264: 289-291.
  • Evolution
    Kornfield, I.L. and R.K. Koehn. 1975. Genetic variation and evolution in some New World cichlids. Evolution 29:427-437.
  • 1975 Quart. Rev. Biology
    Kornfield, I. 1975. The cichlid fishes of Lake Victoria, East Africa: The biology and evolution of a species flock. Quart. Rev. Biol. 50:487.
  • Quart. Rev. Biology
    Kornfield, I. 1974. Laboratory and Field Investigations in General Ecology. Quart. Rev. Biol. 49:165-166.

Are you Irv Kornfield? If so, log in to edit your profile.

Marine Science

Copyright © 2016 UMaine School of Marine Sciences

Website built by RainStorm Consulting