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Susan Brawley

If you need to locate a faculty member with knowledge about particular functions of SMS (e.g., graduate admissions), you can find them on the list of current faculty committees.

Susan Brawley

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Susan Brawley


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321 Hitchner Hall
Orono, ME  04469-5741


Ph.D. University of California

Research interests



  • Muhlin, J. F., C. R. Engel, R. Stessel, R. A. Weatherbee & S. H. Brawley. 2008. The influence of coastal topography, circulation patterns and rafting in structuring populations of an intertidal alga. Molec. Ecol. 17, 1198-1210.
  • Koester, J., S. H. Brawley, L. Karp-Boss and D. G. Mann. 2007. Sexual reproduction in the marine centric diatom Ditylum brightwellii (Bacillariophyta). Eur. J. Phycol. 42, 351-366.
  • Blouin, N. B., X. G. Fei, J. Peng, C. Yarish, and S. H. Brawley. 2007. Seeding nets with neutral spores of the red alga Porphyra umbilicalis (L.) Kützing for use in integrated multi-trophic aquaculture (IMTA). Aquaculture 270, 77-91.
  • Blouin, N., B. L. Calder, B. Perkins, and S. H. Brawley. 2006. Sensory and fatty acid analyses of
    two Atlantic species of Porphyra (Rhodophyta). J. Appl. Phycol. 18, 79-85.
  • Coleman, M. A. and S.H. Brawley. 2005. Variability in temperature and historical patterns in
    reproduction in the Fucus distichus complex (Heterokontophyta; Phaeophyceae): Implications for speciation and the collection of herbarium specimens. J. Phycol. 41, 1110-1119.
  • Coleman, M. and S. H. Brawley. 2005. Spatial and temporal variability in dispersal and
    population genetic structure of a rockpool alga. Mar. Ecol. Prog. Ser. 300, 63-77.
  • Coleman, M. A. and S. H. Brawley. 2005. Are life history characteristics good predictors of genetic diversity and structure? A case study of the interetidal alga Fucus spiralis (Heterokontophyta: Phaeophyceae). J. Phycol. 41, 753-762.
  • Gordon, R. and S. H. Brawley. 2004. Effects of water motion on propagule release from algae with complex life histories. Mar. Biol. 145, 21-29.
  • Calder, E., M. Bagley, and S. H. Brawley. 2004. National Science Foundation Graduate Teaching Fellows promote food science education in K-12 schools in Maine. J. Food Sci. Ed. 2, 58-60.
  • Li, R. and S. H. Brawley. 2004. Improved survival to heat stress in intertidal embryos simultaneously exposed to hypersalinity and the effect of parental thermal history. Mar. Biol. 144, 205-213.
  • Engel, C., S. H. Brawley, K. J. Edwards and E. Serrão. 2003. Isolation and cross-species
    amplification of microsatellite loci from the fucoid seaweeds Fucus vesiculosus, F. serratus and Ascophyllum nodosum (Heterokontophyta, Fucaceae). Molecular Ecology Notes 3,180-182.
  • Berndt, M. L., J. A. Callow, and S. H. Brawley. 2002. Gamete concentrations and timing and success of fertilization in a rocky shore seaweed. Mar. Ecol. Prog. Ser. 226, 273-285.
  • Speransky, V., S. H. Brawley, and M. E. McCully. 2001. Ion fluxes and modification of the extracellular matrix during gamete release in fucoid algae. J. Phycol. 37, 555-573.
  • Brawley, S.H., and K.R. Robinson. 1985. Cytochalasin treatment disrupts the endogenous currents associated with cell polarization in embryogenesis: studies of the role of F-actin in embryogenesis. J. Cell Biol. 100, 1173-84.
  • Speransky, S. and S. H. Brawley. 2000. Gamete release is increased by calm conditions in the coenocytic green alga Bryopsis (Chlorophyta). J. Phycol. 36, 730-739.
  • Brawley, S.H., L.E. Johnson, G.A. Pearson, V. Speransky, R.L. and E. Serrão 1999. Gamete release at low tide in fucoid algae: Maladaptive or advantageous? Amer. Zool. 39: 218-29.
  • Rosemond, A.D., P.J. Mulholland, and S.H. Brawley. 2000. Seasonally shifting limitation of stream periphyton: Response of algal populations and assemblage biomass and productivity to variation in light, nutrients, and herbivores. Can. J. Fish. Aquat. Sci. 57: 66-75.
  • Serrão, E., L. Alice and S. Brawley. 1999. Evolution of the Fucaceae (Phaeophyceae) inferred from nrDNA-ITS. J. Phycol. 35, 382-394.
  • Johnson, L.E. and S.H. Brawley. 1999. Dispersal and recruitment of a canopy-forming intertidal alga: The relative roles of propagule availability and post-settlement processes. Oecologia 117, 517-26.
  • Serrão, E., S.H. Brawley, J. Hedman, L. Kautsky and G. Samuelsson. 1999. Reproductive success in Fucus vesiculosus (Phaeophyceae) in the Baltic Sea. J. Phycol. 35: 254-269.
  • Brawley, S.H. 1999. Submission and retrieval of an aligned set of nucleic acid sequences.
    J. Phycol. 35, 433-37.
  • Speransky, V. S. Speransky, and S. H. Brawley. 1999. Cryoanalytical studies of freezing damage and recovery in Fucus vesiculosus (Phaeophyceae). J. Phycol. 35, 1264-1275.
  • Pearson, G.A., E. Serrão, and S.H. Brawley. 1998. Control of gamete release in fucoid algae: sensing hydrodynamic conditions via carbon acquisition. Ecology 79, 1725-1739.
  • Li, R., S.H. Brawley and T. Close. 1998. Proteins immunologically related to dehydrins in fucoid algae. J. Phycol. 34, 642-50.
  • Pearson, G. and S.H. Brawley. 1998. A model for signal transduction during gamete release in the fucoid alga Pelvetia compressa. Plant Physiol. 118, 305-13.
  • Rosemond, A. D. and S. H. Brawley. 1996. Species-specific characteristics explain the persistence of Stigeoclonium tenue (Chlorophyta) in a woodland stream. J. Phycol. 32, 54-63.
  • Serrão, E. A., L. Kautsky and S. H. Brawley. 1996. Distributional success of the marine seaweed Fucus vesiculosus the brackish Baltic Sea correlates with the osmotic capabilities of Baltic gametes. Oecologia 107, 1-12.
  • Serrão, E. A., G. Pearson, L. Kautsky and S. H. Brawley. 1996. Successful external fertilization in turbulent environments. Proc. Natl. Acad. Sci. U.S.A. 93, 5286-90.
  • Pearson, G. and S. H. Brawley. 1996. Reproductive ecology of Fucus distichus (Phaeophyceae): An intertidal alga with successful external fertilization. Mar. Ecol. Prog. Ser. 143, 211-23.
  • Committee on Biodiversity in Marine Systems. 1995. Understanding Marine Biodiversity. National Academy Press, Washington, 128 pp.
  • Brawley, S.H. and L.E. Johnson. 1993. Predicting desiccation stress in microscopic organisms: the use of agarose beads to determine evaporation rates within and between intertidal microhabitats. J.Phycol. 29, 528-535.
  • Davison, I.R., L.E. Johnson, and S.H. Brawley. 1993. Sublethal stress in the intertidal zone: tidal emersion inhibits photosynthesis and retards development in embryos of the brown alga Pelvetia fastigiata. Oecologia 96, 483-492.
  • Brawley, S.H. 1992. Mesoherbivores. In: Plant-Animal interactions in the Marine Benthos (D. John, S. Hawkins & J. Price, Ed.), Oxford University Press, Oxford, pp 235-263.
  • Brawley, S.H. 1992. Fertilization in natural populations of the dioecious brown alga Fucus ceranoides L. and the importance of the polyspermy block. Marine Biology. 113, 145-157 
  • Brawley, S.H. and L.E. Johnson. 1992. Gametogenesis, gametes and zygotes: an ecological perspective on sexual reproduction in the algae. British Phycological Journal 27, 233-252.
  • Brawley, S.H. 1991. The fast block against polyspermy in fucoid algae is an electrical block. Develop. Biol. 144, 94-106.
  • Brawley, S.H. and L.E. Johnson. 1991. Survival of fucoid embryos in the intertidal zone depends upon developmental stage and microhabitat. J. Phycol. 27, 179-186.
  • Brawley, S.H. 1990. Polyspermy blocks in fucoid algae and the occurrence of polyspermy in nature. In: Mechanism of Fertilization: Plants to Humans (B.Dale, Ed.). Berlin: Springer- Verlag, pp. 419-431.
  • Brawley, S.H. 1990. The polyspermy block in fucoid algae. In: Cell Walls and Surfaces, Reproduction, Photosynthesis, Experimental Phycology I (W. Wiessner, D.G. robinson, R.C. Starr, Eds.). Berlin: Springer-Verlag, pp. 134-144.
  • Brawley, S.H. and D.M. Roberts. 1989. Calmodulin-binding proteins are developmentally regulated in gametes and embryos of fucoid algae. Develop. Biol. 131, 313-320.
  • Brawley, S.H. and X.G. Fei. 1988. Ecological studies of G. asiatica and G. lemaneiformis in China. Chinese J. Oceanol. Limnol. 6, 22-34.
  • Brawley, S.H. and E. Bell. 1987. Partial activation of Fucus eggs with calcium ionophores and low- sodium seawater. Develop. Biol. 122, 217-226.
  • Brawley, S.H. and X. G. Fei. 1987. Studies of mesoherbivory in aquaria and in an unbarricaded mariculture farm on the Chinese coast. J. Phycol. 23, 614-623.
  • Brawley, S.H. 1987. A sodium-dependent, fast block to polyspermy occurs in eggs of fucoid algae. Develop. Biol. 124, 390-397. (Featured on journal cover.)
  • Allen, N.S. and S.H. Brawley. 1984. Observations of exocytosis in Fucus vesiculosus gametes using video-enhanced light microscopy: a video report. Cell Motility 4, 25-27.
  • Brawley, S.H., D.F. Wetherell, and K.R. Robinson. 1984. Electrical polarity in embryos of wild carrot precedes cotyledon development. Proc. Natl. Acad. Sci. U.S.A. 81, 6064-6067.
  • Brawley, S.H. and J.R. Sears. 1982. Septal plugs in a green alga. Amer. J. Bot. 69, 455-463.
  • Brawley, S.H. and W. H. Adey. 1982. Coralliophila abbreviata: a significant corallivore! Bull. Mar. Sci. 32, 595-599.
  • Sears, J.R. and S.H. Brawley. 1982. Smithsoniella gen.nov., a possible evolutionary link between the multicellular and siphonous habits in the Ulvophyceae, Chlorophyta. Amer. J. Bot. 69, 1450-1461.
  • Brawley, S.H. and W.H. Adey. 1981. The effect of micrograzers on algal community structure in a coral reef microcosm. Marine Biology 61, 167-177.
  • 1981. Micrograzers may affect macroalgal density. Nature 292, 177.

  • Brawley, S.H. and R. Wetherbee. 1981. Algal cytology and ultrastructure. In: Biology of the Seaweeds (Botanical Monograph series) (C. Lobban and M. Wynne, eds.). Oxford: Blackwell Scientific Pub. Ltd., pgs. 248-299.
  • Quatrano, R.S., S.H. Brawley, and W.E. Hogsett. 1979. The control of the polar deposition of a sulfated polysaccharide in Fucus zygotes. In: Determinants of Spatial Organization (S.Subtelny and I. R. Konigsberg, eds.). New York: Academic Press, pgs. 77-96.
  • Brawley, S.H. and R.S. Quatrano. 1979a. Effects of microtubule inhibitors on pronuclear migration and embryogenesis in Fucus distichus (Phaeophyta). J. Phycol. 15, 266-272.

  • 1979b. Sulfation of fucoidin in Fucus embryos. IV. Autoradiographic investigations of fucoidin sulfation and secretion during differentiation and the effect of cytochalasin treatment. Develop. Biol. 73, 193-205.
  • Brawley, S.H., R.S. Quantrano, and R. Wetherbee. 1977. Fine-structural studies of the gametes and embryo of Fucus vesiculosus L. (Phaeophyta), III. Cytokinesis and the multicellular embryo. J. Cell Sci. 244, 275-294.
  • Brawley, S. H. and W. H. Adey. 1977. Territorial behavior of three-spot damselfish (Eupomacentrus planifrons) increases reef algal biomass and productivity. Env. Biol. Fish 2,45-51

  • Brawley, S.H., R. Wetherbee, and R. S. Quatrano, 1976a. Fine-structural studies of the gametes and the embryo of Fucus vesiculosus L. (Phaeophyta). I. Fertilization and pronuclear fusion. J. Cell Sci. 20, 233-254.
  • Fine-structural studies of the gametes and embryo of Fucus vesiculosus L. (Phaeophyta). II. The cytoplasm of the egg and young zygote. J. Cell Sci. 20, 255-271.

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