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Northern
Leopard Frog (Rana pipiens)
Description: The leopard frog is a slender,
medium-sized frog; females range from 5.4 to 9.5 cm (2.1-3.7") snout-vent
length, and males range from 5.2 to 8.2 cm (2-3.2"). Only one other
species of frog in Maine, the pickerel frog, has distinct spots.
The leopard frog has 2 to 3 unevenly spaced rows of irregular oval-shaped
dark spots that pattern the dorsum. The pickerel frog, in contrast,
has 2 parallel rows of evenly spaced, squarish spots and is colored
a bright yellow-orange on the under surfaces of the hind legs. Leopard
frog spots have light borders on a background of green and/or brown
shades. There are 2 spots, one over each eye. When inactive, elongated
spots on the legs form 1 to 4 bands (Gilhen 1984).The undersides
of the frog are white to grayish white and are unmottled. The frog
is also able to darken itself when its surroundings are dark (Fernandez
and Bagnara 1991). Two possible benefits to this adaptive coloration
are that the animal is less likely to be seen by a predator, and
that the ability to absorb the sun's energy increases (dark objects
absorb heat better than lighter objects) thus helping the animal
to warm up. The white undersides are thought to mimic or resemble
the reflective surface of the water. This is especially important
if a frog is to go swimming safely and not be detected by a hungry
fish looking upward for a meal! Occasionally, northern leopard frogs
without spots on their backs are reported (Merrell 1965, 1970),
but none has been reported in Maine.
Blue leopard frogs have been documented in Maine,
as have blue bullfrogs and green frogs (Berns and Uhler 1966).The
blue color occurs when the skin lacks the surface pigment that would
normally absorb blue light (see Black 1967 for summary).Without
the pigment, blue light is reflected, and we see a blue frog instead
of a green frog.
During the breeding season, males can be distinguished
from females by comparing the thumbs; males have enlarged or swollen
digits to allow the male to securely clasp the female in amplexus.
Two external vocal sacs are only visible when males are calling.
Tadpoles of the leopard frog have a combined body
and tail length no greater than 8.4 cm (3.3").The pollywog is dark
brown with fine gold spots dorsally with some aggregating on the
sides. A bronze iridescence overlays a cream-colored ventral surface.
The snout area may have an orange appearance, and the eyes are bronze.
The tail, being a lighter color than the body, is marked with scattered
fine spots and pencilings (Wright 1914).
Voice: Leopard frogs give 3 vocalizations:
the advertisement call, the release call, and the warning cry. Associated
with these vocalizations are various types of chuckles described
as grunts, putts, clicks, burps, and quacks (Schimdt 1968).The advertisement
call used to attract females is commonly described as a long drawn-out
guttural snore lasting up to 3 seconds. The call is imitated as
"ir-a-a-a------a-a-h" followed by several "rah-rah-rahs" if the
male is pursuing a female (Noble and Aronson 1942).The advertisement
call is given while the male is spread-eagled on the water surface,
or sometimes underwater (Cook 1984).The release call is a short
series of trills ("ir-a-a-ah---- ir-a-a-ah---- ir-a-a-ah----") used
when males are clasped by other males or by females that are not
gravid (Noble and Aronson 1942).The warning cry, an untrilled scream,
is given with an open mouth when the frog is startled or captured
by a predator.
Audio: Northern Leopard Frog call (coming soon)
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| Northern Leopard Frog |
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